Alexandra Houssaye
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Publications 
  1. Lefebvre, R., Allain, R; & Houssaye, A. 2023. What’s inside a sauropod limb? First three-dimensional investigation of the limb long bone microanatomy of a sauropod dinosaur, Nigersaurus taqueti (Neosauropoda, Rebbachisauridae) and implications for weight-bearing function. Palaeontology
  2. Pintore, R., Cornette, R., Houssaye, A. & Allain, R.  Femora from an exceptionally large population of coeval ornithomimosaurs yield evidence of sexual dimorphism in extinct theropod dinosaurs. eLife 12, e83413. elifesciences.org/articles/83413
  3. Bader, C, Delapré, A. & Houssaye, A. 2023. Shape variation in the limb long bones of modern elephants reveals adaptations to body-mass and habitat. Journal of Anatomy 242, 806-830. onlinelibrary.wiley.com/doi/10.1111/joa.13827
  4. Cottereau, R., Ortiz, K., Locatelli, Y., Houssaye, A. & Cucchi, T.  2023. Can growth in captivity alter the calcaneal microanatomy of a wild ungulate? PCIArcheology Peer Community Journal, Volume 3, article no. e1. peercommunityjournal.org/item/10.24072/pcjournal.210.pdf
  5. Cadoret, N., Chaves-Jacob, J. & Houssaye, A. Form-function relationship between trabecular bone architecture and biomechanical function in the horse humerus. Procedia CIRP, 2022, vol. 110, p. 299-304. www.sciencedirect.com/science/article/pii/S221282712200837X
  6. Nakajima, Y., Shigeta, Y, Houssaye, A., Zakharov, Y.D., Popov, A.M. & Sander, P.M. 2022. Early Triassic ichthyopterygian fossils from the Russian Far East. Scientific Reports 12, 5546. www.nature.com/articles/s41598-022-09481-6#citeas
  7. Lefebvre, R., Houssaye, A., Mallison, H., Cornette, R. & Allain, R. 2022. A path to gigantism: Three-dimensional study of the sauropodomorph limb long bone shape variation in the context of the emergence of the sauropod bauplan. Journal of Anatomy 241: 297-336. onlinelibrary.wiley.com/doi/full/10.1111/joa.13646
  8. Bader, C., Böhmer, C., Abou, M. & Houssaye, A. 2022. How does bone microanatomy and musculature covary? An investigation in the forelimb of two species of martens (Martes foina, Martes martes). Journal of Anatomy 241: 145-167. onlinelibrary.wiley.com/doi/full/10.1111/joa.13645
  9. Mallet C., Billet G., Cornette R. & Houssaye A. 2022. Adaptation to graviportality in Rhinocerotoidea? An investigation through the long bone shape variation in their hind limb. Zoological Journal of the Linnean Society 196: 1235–1271. hal.science/hal-03611156v1
  10. Houssaye, A. 2022. Evolution: Back to heavy bones in salty seas. Current Biology 32, 248-255.e2 hal.science/hal-03820094v1
  11. Pintore, R., Houssaye, A., Nesbitt, S.J. & Hutchinson, J.R. In press. Femoral specializations to locomotor habits in early archosauriforms. Journal of Anatomy 240: 867-892 onlinelibrary.wiley.com/doi/full/10.1111/joa.13598
  12. Mallet C., Houssaye A., Cornette R., Billet G. 2021. Long bone shape variation in the forelimb of Rhinocerotoidea – Relation with size, body mass and body proportions. Zoological Journal of the Linnean Society 196: 1201–1234. hal.science/hal-03611154
  13. Houssaye, A., Perthuis, A. de. & Houée, G. 2021. Sesamoid bones also show functional adaptation in their microanatomy - the example of the patella in Perissodactyla. Journal of Anatomy 240: 50-65. hal.science/hal-03357183v1
  14. Etienne, C., Houssaye, A. & Hutchinson, J. 2021.  Limb myology and muscle architecture of the Indian rhinoceros Rhinoceros unicornis and the white rhinoceros Ceratotherium simum (Mammalia: Rhinocerotidae). PeerJ. peerj.com/articles/11314/
  15. Houssaye, A., Canoville, A. & Davesne, D. 2021. A Methodological Renaissance to Advance on Perennial Issues. In Vertebrate Skeletal Histology and Paleohistology. Eds. V. de Buffrénil, A. de Ricqlès, L. Zylberberg and K. Padian. Boca Raton, FL, USA: CRC Press. Chapter 40.
  16. Houssaye, A. & Buffrénil, V. 2021. Bone histology and the adaptation to aquatic life in tetrapods. In Vertebrate Skeletal Histology and Paleohistology. Eds. V. de Buffrénil, A. de Ricqlès, L. Zylberberg and K. Padian. Boca Raton, FL, USA: CRC Press. Chapter 36.
  17. Scheyer, T, Houssaye, A. & Klein, N. 2021. Sauropterygia: Nothosauria and Pachypleurosauria. In Vertebrate Skeletal Histology and Paleohistology. Eds. V. de Buffrénil, A. de Ricqlès, L. Zylberberg and K. Padian. Boca Raton, FL, USA: CRC Press. Chapter 22.
  18. Buffrénil, V. de & Houssaye, A. 2021. Lepidosauria.In Vertebrate Skeletal Histology and Paleohistology. Eds. V. de Buffrénil, A. de Ricqlès, L. Zylberberg and K. Padian. Boca Raton, FL, USA: CRC Press. Chapter 20.
  19. Houssaye, A., Martin, F., Boisserie JR. & Lihoreau, F. 2021. Paleoecological inferences from long bone microanatomical specializations in Hippopotamoidea (Mammalia, Artiodactyla). Journal of Mammalian Evolution link.springer.com/article/10.1007/s10914-021-09536-  Access: hal.archives-ouvertes.fr/hal-03170283/document
  20. Etienne C., Filippo A., Cornette R. & Houssaye A. 2021. Effect of mass and habitat on the shape of limb long bones: a morpho-functional investigation on Bovidae (Mammalia: Cetartiodactyla). Journal of Anatomy. onlinelibrary.wiley.com/doi/10.1111/joa.13359  Access: hal.archives-ouvertes.fr/hal-03159183/document
  21. Pintore, R., Delapré, A., Lefebvre, R., Botton-Divet, L., Houssaye, A. & Cornette, R. In press. The potential and limits of Thin-Plate Splines retrodeformation on asymmetrical objects: Simulation of taphonomical deformations and application on a fossil sample of limb long bones. CR Palevol.
  22. Lefebvre, R., Allain, R, Houssaye, A. & Cornette R. 2020. Disentangling biological variability and taphonomy: shape analysis of the limb long bones of the sauropodomorph dinosaur Plateosaurus. PeerJ 8:e9359. peerj.com/articles/9359/
  23. Mallet, C., Billet, G, Houssaye, A. & Cornette, R. 2020. A first glimpse at the influence of body mass in the morphological integration of the limb long bones: an investigation in modern rhinoceroses. Journal of Anatomy 237: 704-726 onlinelibrary.wiley.com/doi/abs/10.1111/joa.13232 Access: hal.archives-ouvertes.fr/hal-02869326/document
  24. Etienne, C., Mallet, C., Cornette, R. & Houssaye, A. 2020. Influence of mass on tarsus shape variation: a morphometrical investigation among Rhinocerotidae (Mammalia: Perissodactyla). Biological Journal of the Linnean Society 129:950-974. academic.oup.com/biolinnean/article-abstract/129/4/950/5727857?redirectedFrom=fulltext&casa_token=xjP5BjmEqW0AAAAA:mAZRlfCnpH1yyC1MVS1twknL_IO9S79KwtPIcsYXroi695ZeD0Mu5PucbQC3QFnI8RIvZ8d6lkdkQQ   Access: hal.archives-ouvertes.fr/hal-02566044/document
  25. Houssaye, A. & Prévoteau, J. 2019. What about limb long bone nutrient canal(s)? – A 3D investigation in mammals. Journal of Anatomy 236: 510-521. onlinelibrary.wiley.com/doi/abs/10.1111/joa.13121 Access: hal.archives-ouvertes.fr/hal-02565940/document
  26. Folie, A., Buffetaut, E., Bardet, N., Houssaye, A., Gheerbrant, E. & Laurin, M. 2019. Paléobiologie et paléobiogéographie des amphibiens et reptiles : un hommage à Jean-Claude Rage – 1re partie. CR Palevol 18:693–697. www.sciencedirect.com/science/article/pii/S1631068319301885  Access:  hal.sorbonne-universite.fr/hal-02451171/document
  27. Díaz-Berenguer, E., Houssaye, A., Badiola, A. & Canudo, J. I. 2019. The hind limbs of Sobrarbesiren cardieli (Eocene,Northeastern Spain) and new insights into the locomotion capabilities of the quadrupedal sirenians. J Mammal Evol. link.springer.com/article/10.1007/s10914-019-09482-9  Access: hal.archives-ouvertes.fr/hal-02351001/document
  28. Mallet C., Cornette R., Billet G., Houssaye A. 2019. Interspecific variation in the limb long bones among modern rhinoceroses - extent and drivers. PeerJ 7 e7647  peerj.com/articles/7647/
  29. Houssaye, A., Herrel, A., Boistel R. & Rage, J.C. 2019. Adaptation of the vertebral inner structure to an aquatic life in snakes: pachyophiid peculiarities in comparison to extant and extinct forms. CR Palevol 18:783-799. www.sciencedirect.com/science/article/pii/S1631068319301204  Access: hal.sorbonne-universite.fr/hal-02430856/document
  30. Allemand, R., Houssaye, A., Bardet, N. & Vincent, P. 2019. Endocranial anatomy of plesiosaurians (Reptilia, Plesiosauria) from the Late Cretaceous (Turonian) of Goulmima (Southern Morocco). Journal of Vertebrate Paleontology 39 e1595636 www.tandfonline.com/doi/abs/10.1080/02724634.2019.1595636?journalCode=ujvp20  Access: hal.archives-ouvertes.fr/hal-02354078/document
  31. MacLatchy, L., Smith, T., Houssaye, A., Olejniczak, A. & Rossie, J. 2019. New hominoid fossils from Moroto II, Uganda and their bearing on the taxonomic and adaptive status of Morotopithecus bishopi. Journal of Human Evolution 132: 227-246. www.sciencedirect.com/science/article/abs/pii/S0047248418301568
  32. Klein, N., Canoville, A. & Houssaye, A. 2019. Microstructure of vertebrae, ribs, and gastralia of Triassic sauropterygians
    – New insights into the microanatomical processes involved in aquatic adaptations of marine reptiles. Anatomical Record 302: 1770-1791. anatomypubs.onlinelibrary.wiley.com/doi/full/10.1002/ar.24140
  33. Smith, T., Houssaye, A., Kullmer, O., Le Cabec, A., Olejniczak, A., Schrenk, F., de Vos, J. & Tafforeau, P. 2018. Disentangling Isolated Dental Remains of Asian Pleistocene Hominins and Hominoids. PlosOne 13 e0204737 journals.plos.org/plosone/article?id=10.1371/journal.pone.0204737
  34. Houssaye, A. & Botton-Divet, L. 2018. From land to water: evolutionary changes in long bone microanatomy of otters (Mammalia: Mustelidae). Biological Journal of the Linnean Society 125: 240-249. academic.oup.com/biolinnean/article-abstract/125/2/240/5079862
  35. Houssaye, A., Taverne, M. & Cornette, R. 2018. 3D Quantitative comparative analysis of long bone diaphysis variations in microanatomy and cross-sectional geometry. Journal of Anatomy 232: 836-849. onlinelibrary.wiley.com/doi/full/10.1111/joa.12783
  36. Botton-Divet, L., Houssaye, A., Herrel, A., Fabre, AC & Cornette R. 2018. Swimmers, diggers, climbers and more, a study of integration across the mustelids'locomotor apparatus (Carnivora: Mustelidae). Evolutionary Biology 45: 182–195. link.springer.com/article/10.1007/s11692-017-9442-7
  37. Houssaye, A., Nakajima, Y. & Sander P.M. 2018. Structural, functional, and physiological signals in ichthyosaur vertebral centrum microanatomy and histology. Geodiversitas  40: 161-170. sciencepress.mnhn.fr/en/periodiques/geodiversitas/40/7
  38. Allemand, R., Bardet, N., Houssaye, A. & Vincent, P. 2018. New plesiosaurian specimens (Reptilia, Plesiosauria) from the Upper Cretaceous (Turonian) of Goulmima (Southern Morocco). Cretaceous Research. 82: 83-98. www.sciencedirect.com/science/article/abs/pii/S0195667117302380
  39. Wintrich, T., Hayashi, S., Houssaye, A., Nakajima, Y., Sander, P. M. 2017. A Triassic plesiosaurian skeleton and bone histology inform on evolution of a unique body plan and survival of end-Triassic extinctions. Science Advances 3(12): e1701144. advances.sciencemag.org/content/3/12/e1701144.full
  40. Allemand, R. Boistel, R., Daghfous, G., Blanchet, Z., Cornette, R., Bardet, N., Vincent, P. & Houssaye, A. 2017. Comparative morphology of snake (Squamata) endocasts: evidence of phylogenetical and ecological signals. Journal of Anatomy 231: 849-868. onlinelibrary.wiley.com/doi/10.1111/joa.12692
  41. Allemand, R., Bardet, N., Houssaye, A. & Vincent, P. 2017. Virtual re-examination of a plesiosaurian specimen (Reptilia, Plesiosauria) from the Late Cretaceous (Turonian) of Goulmima (Southern Morocco) thanks to computed tomography. Journal of Vertebrate Paleontology 37: e1325894. www.tandfonline.com/doi/abs/10.1080/02724634.2017.1325894
  42. Bardet, N., Gheerbrant, E., Noubhani, A., Cappetta, H., Jouve, S., Bourdon, E., Pereda Suberbiola, X., Jalil, N.-E., Vincent, P., Houssaye, A., Solé, F., El Houssaini, K., Adnet, S., Rage, J.-C., Lapparent, F. de, Sudre, J., Bouya, B., Amaghzaz, M. & Meslouh, S. 2017. Les Vertébrés des Phosphates crétacés-paléogènes (70,6 - 46,6 Ma) du Maroc. In: Paléontologie des Vertébrés du Maroc : état des connaissances (Ed. S. Zouhri). Mémoires Société Géologique de France, pp. 351-452. /C:/Users/Alex/AppData/Local/Temp/VetrbrsdesPhosphatesduMaroc.pdf
  43. Olivier, C., Houssaye, A., Jalil, N.E. & Cubo, J. 2017. First paleohistological inference of resting metabolic rate in an extinct synapsid, Moghreberia nmachouensis (Therapsida, Anomodontia). Biological Journal of the Linnean Society 121: 409-419. academic.oup.com/biolinnean/article/121/2/409/2999304
  44. Botton-Divet, L., Cornette R., Houssaye A., Fabre A.-C., Herrel A. 2017. Swimming and running, a study of the convergences in long bone morphology among semi-aquatic mustelids (Carnivora: Mustelidae). Biological Journal of the Linnean Society 121: 38-49. academic.oup.com/biolinnean/article/121/1/38/3060320
  45. Houssaye, A., Sander P.M. & Klein, N. 2016. Adaptive patterns in aquatic amniote bone microanatomy – More complex than previously thought. Integrative & Comparative Biology 56: 1349-1369. academic.oup.com/icb/article/56/6/1349/2410949
  46. Botton-Divet, L., Cornette, R., Fabre, AC., Herrel, A. & Houssaye, A. 2016. Morphological analysis of long bones in semi-aquatic mustelids and their terrestrial relatives. Integrative & Comparative Biology 56: 1298-1309. academic.oup.com/icb/article/56/6/1298/2410955?login=true
  47. Houssaye, A. & Fish, FE. 2016. Functional (secondary) adaptation to an aquatic life in vertebrates: an introduction to the symposium. Integrative & Comparative Biology 56: 1266-1270. academic.oup.com/icb/article/56/6/1266/2647109
  48. Fau, M., Cornette, R. & Houssaye, A. 2016. Photogrammetry for 3D digitizing bones of mounted skeletons – potential and limits. Comptes Rendus Palevol 15: 968-977. www.sciencedirect.com/science/article/pii/S1631068316300732
  49. Klein, N., Sander, PM., Krahl, A., Scheyer, TM. &  Houssaye, A . 2016.  Aquatic adaptations in Nothosaurus spp. (Sauropterygia)—inferences from humeral histology and microanatomy. PlosOne 11(7): e0158448. journals.plos.org/plosone/article?id=10.1371/journal.pone.0158448
  50. Houssaye, A., Fernandez, V. & Billet, G. 2016. Hyperspecialization in some South American endemic ungulates revealed by long bone microstructure. Journal of Mammalian Evolution.23: 221–235. link.springer.com/article/10.1007%2Fs10914-015-9312-y
  51. Pérez-Garcia, A., Ortega, F., Bolet, A., Escaso, F., Houssaye, A., Martinez-Salanova, J., de Miguel Chaves, C., Mocho, P., Narvaez, I., Segura, M., Torices, A., Vidal, D. & Sanz, J.L. 2016. An update on the knowledge of the Spanish upper Campanian vertebrate site of Armuna (Segovia Province). Cretaceous Research 57: 591-623. www.sciencedirect.com/science/article/abs/pii/S0195667115300513
  52. Houssaye, A., Waskow, K., Hayashi, S., Cornette, R., Lee, A.H., Hutchinson, J.R. 2016. Biomechanical evolution of solid bones in large animals: a microanatomical investigation. Biological Journal of the Linnean Society 117(2): 350-371 www.onlinelibrary.wiley.com/doi/abs/10.1111/bij.12660
  53. Botton-Divet, L., Houssaye, A., Herrel, A., Fabre, A-C., Cornette, R. 2015. Tools for quantitative form description; an evaluation of different software packages for semi-landmark analysis. PeerJ 3:e1417 peerj.com/articles/1417/
  54. Garcia, G., Bardet, N., Houssaye, A., Pereda Suberbiola, X. & Valentin, X. 2015. Mosasauroid (Squamata) discovery in the Late Cretaceous (Early Campanian) continental deposits of Villeveyrac-Lolivet, southern France. C.R. Palevol 14: 6-7. www.sciencedirect.com/science/article/pii/S1631068315000664
  55. Houssaye, A., Tafforeau, P., Muizon, C. de & Gingerich, PD. 2015. Transition of Eocene whales from land to sea: evidence from bone microstructure. Plos One 10(2): e0118409. journals.plos.org/plosone/article?id=10.1371/journal.pone.0118409
  56. Ortega, F., Bardet, N., Barroso-Barcenilla, F. , Callapez, P.M., Cambra-Moo, O., Daviero-Gómez, V., Díez Díaz, V., Domingo, L., Elvira, A., Escaso, F., García-Oliva, M., Gómez, B., Houssaye, A., Knoll, F., Marcos-Fernández, F., Martín, M., Mocho, P., Narváez, I., Pérez- García, A., Peyrot, D., Segura, M., Serrano, H., Torices, A., Vidal, D. & Sanz,J.L. 2015. The biota of the Upper Cretaceous site of Lo Hueco (Cuenca, Spain). Journal of Iberian Geology 41 (1): 83-99. /C:/Users/Alex/AppData/Local/Temp/48657-Article%20Text-82905-2-10-20150508.pdf
  57. Smith, T.M., Tafforeau, P., Le Cabec, A., Bonnin, A., Houssaye, A., Pouech, J., Moggi-Cecchi, J., Manthi, F., Ward, C., Makaremi, M. & Menter, C.G. 2015. Dental Ontogeny in Pliocene and Early Pleistocene Hominins. Plos One 10(2): e0118118. journals.plos.org/plosone/article?id=10.1371/journal.pone.0118118
  58. Hontecillas, D., Houssaye, A., Laplana, C., Sevilla Garcia, P., Luis Arsuaga, J., Péez-Gonzalez, A., Baquedano, E. & Knoll, F. 2015. Reworked marine pythonomorph (Reptilia, Squamata) remains in Late Pleistocene cave deposits in central Spain. Cretaceous Research 54: 188-202. www.sciencedirect.com/science/article/abs/pii/S0195667114002407
  59. Klein, N., Houssaye, A., Neenan, J.M. & Scheyer, T.M. 2015. Long bone histology and microanatomy pf Placodontia (Diapsida: Sauropterygia). Contributions to Zoology 84: 59-84. brill.com/view/journals/ctoz/84/1/article-p59_5.xml
  60. Bardet, N., Houssaye, A., Vincent, P., Pereda Suberbiola, X., Amaghzaz, M., Jourani, E. & Meslouh, S. 2015. Palaeoecology of mosasaurids (Squamata) from the Maastrichtian Phosphates of Morocco. Gondwana Research 27(3): 1068-1078. www.sciencedirect.com/science/article/abs/pii/S1342937X1400272X
  61. Houssaye, A. 2014. Bone histology of fossil tetrapods: advancing methods, analysis, and interpretation. Journal of Vertebrate Paleontology 34(5): 1258. www.tandfonline.com/doi/abs/10.1080/02724634.2013.812102
  62. Bardet, N., Falconnet, J.,  Fischer, V., Houssaye, A., Jouve, S., Pereda Suberbiola, X., Pérez-García, A., Rage, J.-C. & Vincent, P. 2014. Mesozoic marine reptile palaeobiogeography in response to drifting plates. Gondwana Research 26:869-887. www.sciencedirect.com/science/article/abs/pii/S1342937X1400183X
  63. Houssaye, A. 2014. Advances in vertebrate paleohistology: recent progresses, discoveries and new approaches. Biological Journal of the Linnean Society 112(4): 645-648. academic.oup.com/biolinnean/article/112/4/645/2415934?casa_token=IaPcjh256okAAAAA:Pm1ZoDkr4pgUFMosIoe7IZG-eWMjSEg8DcMG7jJJ2GZuWUo_ReMkxLcZm9enbMaOvMuGpkexAQB9XA
  64. Houssaye, A., Tafforeau, P. & Herrel, A. 2014. Amniote vertebral microanatomy - what are the major trends? Biological Journal of the Linnean Society 112(4): 735-746. onlinelibrary.wiley.com/doi/abs/10.1111/bij.12311
  65. Nakajima, Y.,Houssaye, A. & Endo, H. 2014. Osteohistology of Utatsusaurus hataii (Reptilia: Ichthyopterygia): Implications for early ichthyosaur biology. Acta Palaeontologica Polonica 59(2): 343-352. bioone.org/journals/Acta-Palaeontologica-Polonica/volume-59/issue-2/app.2012.0045/Osteohistology-of-the-Early-Triassic-Ichthyopterygian-Reptile-Utatsusaurus-hataii/10.4202/app.2012.0045.full
  66. Houssaye, A., Scheyer, T.M., Kolb, C., Fischer, V. & Sander, PM. 2014. A new look at ichthyosaur long bone microanatomy and histology: Implications for their adaptation to an aquatic life. Plos One 9(4): e95637 journals.plos.org/plosone/article?id=10.1371/journal.pone.0095637
  67. Houssaye, A. & Bardet, N. 2013. A baby mosasauroid (Reptilia, Squamata) from the Turonian of Morocco - Tethysaurus 'junior' discovered? Cretaceous Research 46:208-215. www.sciencedirect.com/science/article/abs/pii/S0195667113001419
  68. Houssaye, A., Boistel, R., Böhme, W., & Herrel, A. 2013. Jack of all trades master of all? Snake vertebrae have a generalist inner organization. Naturwissenschaften 100(11):997-1006. link.springer.com/article/10.1007/s00114-013-1102-x
  69. Houssaye, A., Lindgren, J., Pellegrini, R., Lee, A.H., Germain, D. & Polcyn, M.J. 2013. Microanatomical and histological features in the long bones of mosasaurine mosasaurs (Reptilia, Squamata) – Implications for aquatic adaptation and growth rates. Plos One 8(10): e76741. doi:10.1371/journal.pone.0076741. journals.plos.org/plosone/article?id=10.1371/journal.pone.0076741
  70. Hayashi, S., Houssaye, A., Nakajima, Y., Chiba, K., Inuzuka, N., Sawamura, H., Ando, T., Osaki, T. & Kaneko, N. 2013. Bone histology suggests increasing aquatic adaptations in Desmostylia (Mammalia, Afrotheria). Plos One 8(4)e59146 journals.plos.org/plosone/article?id=10.1371/journal.pone.0059146
  71. Houssaye, A., Bardet, B., Ortega, P. & Narvaez, I. 2013. Squamate finding in "Lo Hueco" (upper Campanian-lower Maastrichtian, Cuenca Province, Spain): The second non-marine pythonomorph lizard. Paläontologische Zeitschrift 87(3):415-422. link.springer.com/article/10.1007/s12542-013-0164-6
  72. Vincent, P., Bardet, N., Houssaye, A., Pereda Suberbiola, X., Amaghzaz, M. & Meslouh, S. 2013. New plesiosaur specimens from the Maastrichtian Phosphates of Morocco: implications for niche partitioning in the latest Cretaceous marine top-predators. Gondwana Research 24(2): 796-805. www.sciencedirect.com/science/article/abs/pii/S1342937X12003875
  73. Houssaye, A., Rage, J.-C., Torcida Fernández-Baldor, F., Huerta Hurtado, P., Bardet, N. & Pereda Suberbiola, X. 2013. A new varanoid squamate from the Early Cretaceous of Burgos, Spain. Cretaceous Research 41:127-135. www.sciencedirect.com/science/article/abs/pii/S0195667112001723
  74. Houssaye, A., Rage, J.-C., Bardet, N., Vincent, P., Amaghzaz, M. & Meslouh, S. 2013. New highlights about the enigmatic marine snake Palaeophis maghrebianus (Palaeophiidae; Palaeophiinae) from the Ypresian Phosphates (Middle Eocene) of Morocco. Palaeontology 56(3):647–661. onlinelibrary.wiley.com/doi/full/10.1111/pala.12008
  75. Houssaye, A. 2013. Bone histology of aquatic reptiles: what does it tell us about secondary adaptation to an aquatic life. Biological journal of the Linnean Society 108(1):3-21. onlinelibrary.wiley.com/doi/abs/10.1111/j.1095-8312.2012.02002.x
  76. Houssaye, A. 2013. Paleoecological and morphofunctional interpretation of bone mass increase: an example in Late Cretaceous shallow marine squamates. Biological Reviews 88(1):117-139.  onlinelibrary.wiley.com/doi/abs/10.1111/j.1469-185X.2012.00243.x
  77. Houssaye, A. & Tafforeau, P. 2012. What vertebral microanatomy reveals about the ecology of juvenile mosasaurs (Reptilia, Squamata). Journal of vertebrate Paleontology 32(5):1042-1048. www.tandfonline.com/doi/abs/10.1080/02724634.2012.680999
  78. Houssaye, A. & Bardet, N. 2012. Ribs and vertebrae microanatomical characteristics in hydropelvic mosasauroids. Lethaia 45(2):200-209. onlinelibrary.wiley.com/doi/abs/10.1111/j.1502-3931.2011.00273.x
  79. Houssaye, A., Xu, F., Helfen, L., Buffrénil, V. de, & Tafforeau, P. 2011. Three dimensional pelvis and limb anatomy of the Cenomanian Lebanese hind-limbed snake Eupodophis descouensi (Squamata, Ophidia) revealed by synchrotron-radiation computed laminography. Journal of Vertebrate Paleontology 31(1):2-7. www.tandfonline.com/doi/abs/10.1080/02724634.2011.539650
  80. Houssaye, A., Bardet, N., Rage, J.-C., Pereda Suberbiola, X., Bouya, B., Amaghzaz, M. & Amalik, M. 2011. A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous Phosphates of the margin of the Southern Tethys. Geological Magazine 148(2):237-249. www.cambridge.org/core/journals/geological-magazine/article/abs/div-classtitlea-review-of-span-classitalicpachyvaranus-crassispondylusspan-arambourg-1952-a-pachyostotic-marine-squamate-from-the-latest-cretaceous-phosphates-of-morocco-and-syriadiv/FC73BFC004B783BFAFC7BD9E96E20DF2
  81. Houssaye, A., Mazurier, A., Herrel, A., Volpato, V., Tafforeau, P., Boistel, R. and Buffrénil, V. de. 2010. Vertebral microanatomy in squamates: structure, growth and ecological correlates. Journal of Anatomy 217(6):715-727. onlinelibrary.wiley.com/doi/full/10.1111/j.1469-7580.2010.01307.x
  82. Bardet, N., Pereda Suberbiolab, X., Jouve, S., Bourdon, E., Vincent, P. Houssaye, A., Rage, J.-C., Jalil, N.-E., Bouya, B., and Amaghzaz, M. 2010. Reptilian assemblages from the latest Cretaceous - Palaeogene phosphates of Morocco: from Arambourg to present time. Historical Biology 22(1-3):186-199. www.tandfonline.com/doi/abs/10.1080/08912961003754945
  83. Houssaye, A. 2010. Rediscovery and description of the second specimen of the hind-limbed snake Pachyophis woodwardi Nopcsa, 1923 (Squamata, Ophidia) from the Cenomanian of Bosnia Herzegovina. Journal of Vertebrate Paleontology 30(1):1-4. www.tandfonline.com/doi/full/10.1080/02724630903416043
  84. Houssaye, A. 2010. A new aquatic pythonomorph (Reptilia, Squamata) from the Turonian (Late Cretaceous) of France. Comptes Rendus Palevol 9:39-45. www.sciencedirect.com/science/article/pii/S1631068309001304
  85. Houssaye, A. 2009. "Pachyostosis" in aquatic amniotes: a review. Integrative Zoology 4:325-340. onlinelibrary.wiley.com/doi/abs/10.1111/j.1749-4877.2009.00146.x
  86. Bardet, N., Houssaye, A., Pereda Suberbiola, X., & Rage, J.-C. 2008. The Cenomanian-Turonian (Late Cretaceous) radiation of marine squamates (Reptilia): the role of the Mediterranean Tethys. Bulletin de la Société Géologique de France 179(6):605-622. pubs.geoscienceworld.org/sgf/bsgf/article-abstract/179/6/605/88534/The-Cenomanian-Turonian-late-Cretaceous-radiation?redirectedFrom=fulltext
  87. Buffrénil, V. de, Houssaye, A., & Böhme, W. 2008. Bone vascular supply in monitor lizards (Squamata: Varanidae): Influence of size, growth, and phylogeny. Journal of Morphology 269:533-543. onlinelibrary.wiley.com/doi/abs/10.1002/jmor.10604
  88. Houssaye, A., Buffrénil, V. de, Rage, J.-C., & Bardet, N. 2008. An analysis of vertebral "pachyostosis" in Carentonosaurus mineaui (Mosasauroidea, Squamata) from the Cenomanian (early Late Cretaceous) of France, with comments on its phylogenetic and functional significance. Journal of Vertebrate Paleontology 28(3):685-691. www.tandfonline.com/doi/abs/10.1671/0272-4634%282008%2928%5B685%3AAAOVPI%5D2.0.CO%3B2
  89. Houssaye, A. 2008. A preliminary report on the evolution of the vertebral microanatomy within mosasauroids (Reptilia, Squamata). In: Everhart M.J., ed. Proceedings of the Second Mosasaur Meeting. Fort Hays State University, Hays, Ka, 81-89. www.researchgate.net/publication/330103642_A_PRELIMINARY_REPORT_ON_THE_EVOLUTION_OF_THE_VERTEBRAL_MICROANATOMY_WITHIN_MOSASAUROIDS_REPTILIA_SQUAMATA
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